Cyrtandra lydgatei

Hillebr. (1888)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Gesneriaceae Genus: Cyrtandra

ha‘iwale [haiwale], kanawao ke‘oke‘o [kanawao keokeo]

Description

Key Characters:

Growth Form: Shrubs 1–2.5 m tall.

Stems: Stems unbranched to few–branched, rounded to angular, pubescent with multicellular hairs or glabrate.

Roots:

Leaves: Leaves simple. Opposite, borne on upper 3–6 nodes. Those leaves of a pair unequal, symmetrical, broadly elliptic to suborbicular, 13–23 cm long, (4.5–)7–10(–14) cm wide. Apex acuminate. Base cuneate or blade abruptly constricted to a long-attenuate base. Upper surfaces sparsely hirsute; lower surfaces pale, sparsely puberulent or glabrate; thin, chartaceous, often flaccid. Margins coarsely and sometimes irregularly serrate. Petioles 3–9 cm long, sparsely to moderately villous. Stipules absent.

Flowers: Flowers (2)3–7 in relatively dense umbelliform cymes arising in the leaf axils, moderately villous throughout, peduncles slender, 5–14 mm long, pedicels slender, unequal in length, 5–16 mm long, bracts lanceolate to elliptic, distinct or occasionally connate, 6–8 mm long. Flowers bisexual (perfect). Calyx 2–5(6)-lobed, actinomorphic or nearly so, green, 9–15 mm long, cleft nearly to base, the lobes linear to linear–spatulate, moderately villous, inner surface glabrous or glabrous in the lower ⅓–1⁄2 and moderately villous in upper part of lobes, deciduous or persistent in fruit. Corolla usually bilabiate, white, tube cylindrical, curved, ca. 8–10 mm long, villous in upper 1⁄2, the lobes ca. 2 mm long in dry specimens, upper lip 2-lobed, lower lip 3-lobed, the lobes imbricate in bud. Stamens 5, inserted about halfway up corolla tube, the 2 upper Stamens fertile, others staminodial, +/– with abortive anthers; fertile anthers coherent positioned in throat of corolla tube, dithecal, opening by longitudinal slits or occasionally by apical pores. Ovary superior, oblong–ovoid, 2-carpellate, 1-celled, placentation parietal, placentas 2, ± intruded, rarely joined in center and dividing ovary into 2 cells; ovules numerous, anatropous, glabrous, apex rounded or with a stylar beak; style 3–4 mm long, glabrous; stigma capitate, 2-lobed, lobes usually elliptic.

Fruit: Fruit a fleshy or firm berry; white; ovoid; 1.3–1.6 cm long; often tipped by the persistent stylar beak. Seeds unknown.

Ploidy:

Habitat: Occurring in wet forest.

Elevation Range: 500–900 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Molokai Endemic
Lana'i Endemic
Maui Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Fl. Hawaiian Isl.: 335 (1888)

Other References

Wagner et al. 1990:767 (Mo, L. WM)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Cyrtandra lydgatei Hillebr. w maui, lahaina dist, kauaula valley, upper s fork below helu PRESERVED_SPECIMEN k. r. wood, s. p. perlman & r. hobdy 3956 Maui US 1/18/1995
2 Cyrtandra lydgatei Hillebr. W Maui, Lahaina Dist, Kauaula Valley, upper S fork below Helu PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P.; Hobdy, R. 3956 Maui US 1/18/1995
3 Cyrtandra lydgatei Hillebr. W Maui, Kauaula velley, back of valley on W side. PRESERVED_SPECIMEN S. P. Perlman & H. Oppenheimer 21289 Maui US 12/3/2008
4 Cyrtandra lydgatei Hillebr. kalawao district, upper wailau valley; olona gulch. PRESERVED_SPECIMEN k. r. wood 1266 Hawaii US 10/2/1991
5 Cyrtandra lydgatei Hillebr. kaloe PRESERVED_SPECIMEN c. n. forbes 31.mo Molokai US
6 Cyrtandra lydgatei Hillebr. Kalawao District, upper Wailau Valley; Olona Gulch. PRESERVED_SPECIMEN Wood, K. R. 1266 Molokai US 10/2/1991
7 Cyrtandra lydgatei Hillebr. Kaloe PRESERVED_SPECIMEN Forbes, C. N. 31.Mo Molokai US 6/12/2020
8 Cyrtandra lydgatei Hillebr. kalae PRESERVED_SPECIMEN forbes, c.n. collector number: 31.mo Molokai BISH 6/1/1912
9 Cyrtandra lydgatei Hillebr. maunalei, head, middle valley PRESERVED_SPECIMEN munro, g.c. collector number: 489 Molokai BISH 1890-06-14
10 Cyrtandra lydgatei Hillebr. wailau valley, east bank of wailau stream, in side gully In side gully PRESERVED_SPECIMEN fosberg, f.r. collector number: 13470 Molokai BISH 12/28/1936
11 Cyrtandra lydgatei Hillebr. USA, Hawaiian Islands, Maui, west Maui, Wailuku District West Maui Natural Area Reserve, Kahakuloa Section, western Kahakuloa Stream headwaters Rare, herbs in understory along perennial; stream. About 5-10 plant observed in tis location with C. hashimotoi, C. Paludosa, C. grayana, C. platyphylla and C. hawaiensis. Metrosideros, Cheirodendron, Dicranopteris, Diplopterygium, Melicope, Coprosma, Kadua, Broussaisia, Dubautia, Pipturus, Touchardia, Urera, Cyrtandra, Clermontia, Cyanea, Perrottentia, Scaevola, Gunnera, Cibotium, Sadleria, Diplazium, Athyrium, Cyclosorus, Smilax, Deparia, Carex, Astelia, Eragrostis, Peperomia, Pilea, Deschampsia, Clidemia, Ticbouchina, Ageratina. PRESERVED_SPECIMEN Oppenheimer, H.L. H111304 Maui BISH 11/6/2013
12 Cyrtandra lydgatei Hillebr. Kalawao District; Upper Wailau Valley; Olona Gulch Alien Veg. & Metrosideros-Dicranopteris lowland wet forest with... Freycinetia, Broussaisia, Antidesma, Cibotium, Psychotria, Hedyotis, Clermontia, Ilex, Myrsine, Cyrtandra, Artemisia, Touchardia. Weeds... Clidemia, Kalachoe, Nephrolepis, Rubus rosifolius, Psidium. PRESERVED_SPECIMEN Wood, K.R. 1266 Molokai BISH 10/2/1991
13 Cyrtandra lydgatei Hillebr. Wailau Valley, Hanuu [sp?] Pali near summit Moss forest of Metrosideros, Gouldia, Broussaisia. PRESERVED_SPECIMEN Nagata, K.M. 1144 Molokai BISH 2/23/1973
14 Cyrtandra lydgatei Hillebr. west maui, kaanapali, honokawai gulch PRESERVED_SPECIMEN hillebrand, w.b. lydgate, j.m. collector number: s.n. Molokai BISH 1870-08-08
15 Cyrtandra lydgatei Hillebr. Pukoo PRESERVED_SPECIMEN Forbes, C.N. 279.MO Molokai BISH 7/1/1912
16 Cyrtandra lydgatei Hillebr. West Maui, Wailuku District, Iao Valley, Poohahoahoa Stream, back of valley. Vegetation: Metrosideros, Cheirodendron, Perrottetia, Psychotria, Kadua, Pipturus, Cyrtandra ssp., Broussaisia, Casuarina, Psidium, Boehmeria, Cyclosorus, Crigeron, Ageratina, Diplazium, Phyllostegia, Cyanea, Tibouchina, Amauropeta, Peperomia, Touchardia, Sadleria, Selaginella, Buddleia, Dubautia, Rubus, Blechnum, Tectaria, Dryopteris, Microlepia. PRESERVED_SPECIMEN Oppenheimer, H.L. H71405 Maui BISH 7/24/2014
17 Cyrtandra lydgatei Hillebr. Honokohau drainage basin PRESERVED_SPECIMEN Forbes, C.N. 485.M Maui BISH 9/25/1917
18 Cyrtandra lydgatei Hillebr. east maui, honomanu gulch PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Maui BISH 5/1/1911
19 Cyrtandra lydgatei Hillebr. honokohau drainage basin Growing alongside a road in a pineapple field at the top of the gulch. PRESERVED_SPECIMEN forbes, c.n. collector number: 485.m Maui BISH 9/25/1917
20 Cyrtandra lydgatei Hillebr. w maui, kauaula velley, back of valley on w side. PRESERVED_SPECIMEN s. p. perlman & h. oppenheimer 21289 Maui US 12/3/2008
21 Cyrtandra lydgatei Hillebr. pukoo PRESERVED_SPECIMEN forbes, c.n. collector number: 279.mo Molokai BISH 7/1/1912
22 Cyrtandra lydgatei Hillebr. Mapulehu Valley PRESERVED_SPECIMEN Forbes, C.N. 310.MO Molokai BISH 7/1/1912
23 Cyrtandra lydgatei Hillebr. Maunalei Gulch, middle valley, at head of gulch PRESERVED_SPECIMEN Munro, G.C. 683 Lanai BISH 1890-00-00
24 Cyrtandra lydgatei Hillebr. Maunalei, head, middle valley PRESERVED_SPECIMEN Munro, G.C. 489 Lanai BISH 1890-00-00
25 Cyrtandra lydgatei Hillebr. mapulehu valley Wet canyon bottom. PRESERVED_SPECIMEN st.john, h. baker, r.j.; coulter, j.w.; yuncker, t.g. collector number: 12752 Molokai BISH 12/28/1932
26 Cyrtandra lydgatei Hillebr. kauaula valley, back of valley on w side In metrosideros polymorpha-Cheirodendron trigynum wet forest with Pipturus albidus, Boehmeria grandis, Sadleria cyatheoides, Dubautia plantaginea, Machaerina angustifolia, Perrottetia sandwicensis, Cyrtandra grayi, Touchardia latifolia, Urera glabra, Coprosma pubens, Broussaisia arguta, Dodonaea viscosa, Cyrtandra hashimotoi. PRESERVED_SPECIMEN perlman, s.p. et al. collector number: 21289 Maui BISH 12/3/2008
27 Cyrtandra lydgatei Hillebr. West Maui, Wailuku District, 'Iao Valley, Nakalaloa Stream Grows alongside C. hawaiiensis, C. hashimotoi, C. platyphylla and C. grayi; no hybrids observed Vegetation: Metrosideros, Cheirodendron, Cyanea, Clermontia, Antidesma, Xylosma, Freycinetia, Urera, Touchardia, Pipturus, Boehmeria, Psychotria, Cyrtandra, Gunnera, Bidens PRESERVED_SPECIMEN Oppenheimer, H.L. H91114 Maui BISH 9/15/2011
28 Cyrtandra lydgatei Hillebr. maunalei gulch, middle valley, at head of gulch PRESERVED_SPECIMEN munro, g.c. collector number: 683 Lanai BISH 1890-06-14
29 Cyrtandra lydgatei Hillebr. mapulehu valley Acacia-Metrosideros mesic forest. W/Alphitonia, Dodonaea, Santalum freycinetianum, Antidesma, Nestegis, Leptecophylla, Melicope ovata, M. barbigera, Pouteria, terrestrial cover of Carex wahuensis, C. meyenii, Gahnia beecheyi, Dianell PRESERVED_SPECIMEN forbes, c.n. collector number: 310.mo Kauai BISH 7/1/1912
30 Cyrtandra lydgatei Hillebr. Wailau Valley, Hanuu [sp?] Pali near summit Moss forest of Metrosideros, Goouldia, Broussaisia [? And ferns?] PRESERVED_SPECIMEN Nagata, K.M. 1144 Molokai BISH 2/23/1973
31 Cyrtandra lydgatei Hillebr. W Maui, Honokahua Valley, along flank of a streambed near headwaters region. In vicinity of Transect Station 2.20 Assoc. vegetation: Gunnera petaloidea, Cheirodendron trigynum, Nothocestrum longifolium, Pipturus albidus, Urera glabra, Cyanea macrostegia, Cyrtandra grayana, C. hawaiiensis, Diplazium sandwichianum. PRESERVED_SPECIMEN Meidell, J.S. 162 Maui BISH 1/7/1997
32 Cyrtandra lydgatei Hillebr. molokai PRESERVED_SPECIMEN nagata, k.m. collector number: 1144 Molokai BISH 8/8/1973
33 Cyrtandra lydgatei Hillebr. Kauaula valley, back of valley on W side In metrosideros polymorpha-Cheirodendron trigynum wet forest with Pipturus albidus, Boehmeria grandis, Sadleria cyatheoides, Dubautia plantaginea, Machaerina angustifolia, Perrottetia sandwicensis, Cyrtandra grayi, Touchardia latifolia, Urera glabra, Coprosma pubens, Broussaisia arguta, Dodonaea viscosa, Cyrtandra hashimotoi. PRESERVED_SPECIMEN Perlman, S. 21289 Maui BISH 12/3/2008
34 Cyrtandra lydgatei Hillebr. Kalae PRESERVED_SPECIMEN Rock, J.F.C. s.n. Molokai BISH 5/25/1918
35 Cyrtandra lydgatei Hillebr. Wailau Valley, east bank of Wailau Stream, in side gully In side gully PRESERVED_SPECIMEN Fosberg, F.R. 13470 Molokai BISH 12/28/1936
36 Cyrtandra lydgatei Hillebr. Kahuaawi Gulch, south of Kapuna Springs access road. Vegetation: Metrosideros, Dodonaea, Perrottetia, Broussaisia, Cyrtandra, Clermontia, Cibotium, Dicranopteris, Microlwepia, Pouteria, Ilex, Psychotria, Pisonia, Charpentiera, Melicope, Peperomia, Pneumatopertis. PRESERVED_SPECIMEN Oppenheimer, H.L. H31102 Molokai BISH 3/9/2011
37 Cyrtandra lydgatei Hillebr. hanuu pali near summit. wailau valley Moss forest of Metrosideros, Gouldia, Broussaisia and ferns. PRESERVED_SPECIMEN nagata, k.m. collector number: 1144 Molokai BISH 2/23/1973
38 Cyrtandra lydgatei Hillebr. Kaluaaha, west fork Along perennail stream. Vegetation: Metrosideros, Cheirodendron, Dicranopteris, Diplopterygium, Clidemia, Cyrtandra, Broussaisia, Cibotium, Vandenboschia, Asplenium, Dryopteris, Clermontia, Carex, Athyrium, Kadua, Psidium, Youngia, Ageratum, Ageratina, Diplazium, Peperomia PRESERVED_SPECIMEN Oppenheimer, H.L. H121410 Molokai BISH 12/5/2014
39 Cyrtandra lydgatei Hillebr. waikamoi gulch PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Maui BISH 8/8/1918
40 Cyrtandra lydgatei Hillebr. wailau valley, hanuu pali near summit Moss forest of Metrosideros, Gouldia, Broussaisia. PRESERVED_SPECIMEN nagata, k.m. collector number: 1144 Molokai BISH 2/23/1973
41 Cyrtandra lydgatei Hillebr. east maui, honomanu trail PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Maui BISH 5/1/1911
42 Cyrtandra lydgatei Hillebr. Waikamoi Gulch PRESERVED_SPECIMEN Rock, J.F.C. s.n. Maui BISH 8/1/1918
43 Cyrtandra lydgatei Hillebr. West Maui, Kaanapali, Honokawai Gulch PRESERVED_SPECIMEN Hillebrand, W.B. s.n. Maui BISH 1870-00-00
44 Cyrtandra lydgatei Hillebr. East Maui, Honomanu Gulch PRESERVED_SPECIMEN Rock, J.F.C. s.n. Maui BISH 5/1/1911
45 Cyrtandra lydgatei Hillebr. kalae PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Oahu BISH 5/25/1918
46 Cyrtandra lydgatei Hillebr. Halawa PRESERVED_SPECIMEN Rock, J.F.C. s.n. Molokai BISH 4/1/1910
47 Cyrtandra lydgatei Hillebr. Kalae PRESERVED_SPECIMEN Forbes, C.N. 31.Mo Molokai BISH 6/1/1912
48 Cyrtandra lydgatei Hillebr. Mapulehu Valley Wet canyon bottom. PRESERVED_SPECIMEN St. John, H. 12752 Molokai BISH 12/28/1932
49 Cyrtandra lydgatei Hillebr. Halawa PRESERVED_SPECIMEN Rock, J.F.C. Collector Number: s.n. Molokai BISH 4/1/1910
50 Cyrtandra lydgatei Hillebr. w maui, honolua valley, along rocky banks of intermittent watercourse under Eupatorium on rocky, wet bank PRESERVED_SPECIMEN oppenheimer, h.l. collector number: 191 Oahu BISH 4/23/1997
51 Cyrtandra lydgatei Hillebr. Wailau Valley, Hanuu [sp?] Pali near summit Moss forest of Metrosideros, Gouldia, Broussaisia and ferns. PRESERVED_SPECIMEN Nagata, K.M. 1144 Molokai BISH 2/23/1973
52 Cyrtandra lydgatei Hillebr. W Maui, Honolua Valley, along rocky banks of intermittent watercourse Assoc. vegetation: Touchardia latifolia, Thelypteris cyatheoides, Diplazium sandwichianum, Cyanea scabra, Coprosma pubens, Dicranopteris linearis, Cyrtandra hawaiiensis, Hedyotis hillebrandii, Sadleria souleyetiana, Rubus rosifolius. PRESERVED_SPECIMEN Oppenheimer, H.L. 191 Maui BISH 4/23/1997
53 Cyrtandra lydgatei Hillebr. w maui, honokahua valley, along flank of a streambed near headwaters region. in vicinity of transect station 2.20 Assoc. vegetation: Gunnera petaloidea, Cheirodendron trigynum, Nothocestrum longifolium, Pipturus albidus, Urera glabra, Cyanea macrostegia, Cyrtandra grayana, C. hawaiiensis, Diplazium sandwichianum. PRESERVED_SPECIMEN meidell, j.s. oppenheimer, h.l. collector number: 162 Maui BISH 1/7/1997
54 Cyrtandra lydgatei Hillebr. kalawao district; upper wailau valley; olona gulch On ash on vertical E-facing rock wall in rocky, grass-Pteridium talus slope community PRESERVED_SPECIMEN wood, k.r. perlman, s. collector number: 1266 Maui BISH 10/2/1991
55 Cyrtandra lydgatei Hillebr. halawa PRESERVED_SPECIMEN rock, j.f.c. collector number: s.n. Molokai BISH 4/1/1910
56 Cyrtandra lydgatei Hillebr. W Maui, Lahaina District, Kaua`ula Valley Local and rare in dense shade near streams. Vegetation: Metrosideros, Dodonaea, Coprosma, Xylosma, Dubautia, Touchardia, Pipturus, Boehmeria, Urera, Clermontia, Cyanea, Lobelia, Perrottetia, Myrsine, Sadleria, Deparia, Diplazium, C. platyphylla, C. grayi, C. paludosa, C. hashimotoi, C. grayana, Freycinetia, Peperomia, Kadua. PRESERVED_SPECIMEN Oppenheimer, H.L. H120804 Maui BISH 12/2/2008
57 Cyrtandra lydgatei Hillebr. West Maui, Lahaina District, Kauaula Valley, south fork, back of valley below pali. On boulder strewn tributary. Vegetation: Metrosideros, Cheirodendron, Coprosma, Dodonaea, Touchardia, Pipturus, Boehmeria, Perrottetia, Xylosma, Antidesma, Clermontia, Cyrtandra spp. Bidens, Cibotium, Sadleria, Diplazium, Peperomia, Dicranopteris, Ageratina, Tibouchina, Rubus. PRESERVED_SPECIMEN Oppenheimer, H.L. H11202 Maui BISH 1/5/2012
58 Cyrtandra lydgatei Hillebr. in between kalac and mupulehn? PRESERVED_SPECIMEN w. hillebrand GH
59 Cyrtandra lydgatei Hillebr. maui. sandwich ins. [=hawaiian islands. lat/long is for maui island.] PRESERVED_SPECIMEN hillebrand, w.b. s.n. MEL
60 Cyrtandra lydgatei Hillebr. Mapulehu; upper east side (Wailau Trail to Lae O Kapuna) summit ridge. Elev. 2400--3000 ft. Metrosideros Montane Wet Forest with Cibotium understory and headwaters of Kapuna Gulch with narrow gulches of Metrosideros-Antidesma Lowland Wet Forest. PRESERVED_SPECIMEN K. R. Wood 7178 Molokai PTBG 2/10/1998
61 Cyrtandra lydgatei Hillebr. Kaluaaha, west fork along perennial stream. PRESERVED_SPECIMEN Hank Oppenheimer H121410 Molokai PTBG 12/5/2014
62 Cyrtandra lydgatei Hillebr. West Maui, Iao Valley, Poohahoahoa Stream, back of valley. PRESERVED_SPECIMEN Hank Oppenheimer H71409 Maui PTBG 7/24/2014
63 Cyrtandra lydgatei Hillebr. West Maui, Kauaula valley, north fork of lower valley. Metrosideros- Dicranopteris linearis wet forest forest. PRESERVED_SPECIMEN Steve Perlman 22906 Maui PTBG 6/6/2012
64 Cyrtandra lydgatei Hillebr. W. Maui, Iao Valley. Nakalaloa Stream. PRESERVED_SPECIMEN Hank Oppenheimer H91114 Maui PTBG 9/15/2011
65 Cyrtandra lydgatei Hillebr. Kauaula valley, back of valley on west side. in Metrosideros polymorpha -Cheirodendron trigynum wet forest. PRESERVED_SPECIMEN Steve Perlman 21289 Maui PTBG 12/3/2008
66 Cyrtandra lydgatei Hillebr. Pelekunu Valley, TNC Preserve survey, transect 10, from Pepeopae to Kaunuohua, along Papaala Pali. In Metrosideros polymorpha montane wet forest. PRESERVED_SPECIMEN Steve Perlman 10399 Molokai PTBG 3/31/1989
67 Cyrtandra lydgatei Hillebr. Wailau valley, north coast, seacliffs of Olokui. In Metrosideros- Dicranopteris linearis wet forest. PRESERVED_SPECIMEN Steve Perlman 17521 Molokai PTBG 4/24/2001
68 Cyrtandra lydgatei Hillebr. Kauaula Valley; upper south fork below Helu; Riparian w/ saturated talus & seeping, vertical basalt walls. Elev. 2500-3500 ft. Metrosideros-Dicranopteris Lowland to Montane Wet Forest. PRESERVED_SPECIMEN K. R. Wood 3956 Maui PTBG 1/18/1995
69 Cyrtandra lydgatei Hillebr. W. Maui. Puu Kukui Watershed Management Area. Along a flank above a stream bed near the headwaters region of Honokahua Valley. In the vicinity of Transect Station 2.20. PRESERVED_SPECIMEN J. Scott Meidell & Hank Oppenheimer 156 Maui PTBG
70 Cyrtandra lydgatei Hillebr. Upper Wailau Valley; Olona Gulch. Alien veg. & Metrosideros-Dicranopteris lowland wet forest. PRESERVED_SPECIMEN K. R. Wood 1266 Molokai PTBG 10/2/1991
71 Cyrtandra lydgatei Hillebr. Within tributary of upper Kahoma Stream, along a dry streambed, approximately 100 m North of USFWS Bird Survey TR34, Station 24 in 5 m . Ohia Montane Wet Forest. PRESERVED_SPECIMEN J. Scott Meidell & Hank Oppenheimer 189 Maui PTBG 3/20/1997
72 Cyrtandra lydgatei Hillebr. Along the rocky banks of an intermittent watercourse of Honolua Valley. PRESERVED_SPECIMEN Hank Oppenheimer 190 Maui PTBG 4/23/1997
73 Cyrtandra lydgatei Hillebr. W. Maui. Kauaula Valley. PRESERVED_SPECIMEN Hank Oppenheimer H120804 Maui PTBG 12/2/2008
74 Cyrtandra lydgatei Hillebr. Honomanu Trail, East Maui. PRESERVED_SPECIMEN J. F. Rock s.n. Maui PTBG 5/1/1911
75 Cyrtandra lydgatei Hillebr. w. maui, puu kukui watershed management area, near region of honokahua valley. PRESERVED_SPECIMEN s. meidel & h. l. oppenheimer 156 Oahu US